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Ecosystem susceptibility to invasion by nonnative species is poorly understood, but evidence is increasing that spatial and temporal variability in resources has large-scale effects. We conducted a study in Artemisia tridentata ecosystems at two Great Basin locations examining differences in resource availability and invasibility of Bromus tectorum over elevation gradients and in response to direct and interacting effects of removal of perennial herbaceous vegetation and fire. We monitored environmental conditions, soil variables, and B. tectorum establishment and reproduction over two years. Soil water (measured as the number of days soil matric potential was >?1.5 MPa) and nitrate availability (measured as micromoles...