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1. Heating and cooling rates of the desert tortoise, Gopherus agassizii, were measured in the field and in the laboratory. 2. Heating rates in the field were up to ten times faster than cooling rates. 3. Under controlled conditions, heating rates equalled rates of cooling. 4. Heart rates during heating were significantly faster than during cooling at any particular body temperature. 5. Subcutaneous neck temperatures did not differ from cloacal temperatures during heating or cooling, suggesting a state of continuous vasodilation. 6. Results are interpreted as suggesting that behavioral postures and activities play a greater role than physiology in the determination of thermal exchange rates of the desert tortoise....